(2021). Intergenerational niche differentiation in filmy ferns (Hymenophyllaceae).

Preprint PDF Code Dataset Project DOI

(2020). A taxonomic and molecular survey of the pteridophytes of the Nectandra Cloud Forest Reserve, Costa Rica. PLoS ONE 15(11): e0241231.

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(2019). Virtual Issue: Ecology and Evolution of Pteridophytes in the Era of Molecular Genetics. J. Plant Res. 132: 719–721.

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(2019). Growth Dynamics of Independent Gametophytes of Pleurosoriopsis makinoi (Polypodiaceae). Bull. Natl. Mus. Nat. Sci., Ser. B 45: 77–86.

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(2018). Keeping an eye on coloration: Ecological correlates of the evolution of pitcher traits in the genus Nepenthes (Caryophyllales). Biological Journal of the Linnean Society 123: 321–337.

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(2017). A plastid phylogeny and character evolution of the Old World fern genus Pyrrosia (Polypodiaceae) with the description of a new genus: Hovenkampia (Polypodiaceae). Molecular Phylogenetics and Evolution 114: 271–294.

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(2017). The Separation of Generations: Biology and Biogeography of Long-Lived Sporophyteless Fern Gametophytes. International Journal of Plant Sciences 178: 1–18.

Open Access

(2017). Life cycle matters: DNA barcoding reveals contrasting community structure between fern sporophytes and gametophytes. Ecological Monographs 87: 278–296.

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(2016). Fern species richness and abundance are indicators of climate change on high-elevation islands: evidence from an elevational gradient on Tahiti (French Polynesia). Climatic Change 138: 143–156.

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(2015). Antrophyum solomonense (Pteridaceae), a New Species from the Solomon Islands, and its Systematic Position Based on Phylogenetic Analysis. Systematic Botany 40: 645–651.

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(2013). A survey of the fern gametophyte flora of Japan: Frequent independent occurrences of noncordiform gametophytes. American Journal of Botany 100: 735–743.

Project Open Access

(2011). Reticulate evolution in the Crepidomanes minutum species complex (Hymenophyllaceae). American Journal of Botany 98: 1782–1800.

Project Open Access

(2011). Pteridophytes of Mo'orea, French Polynesia: additional new records. American Fern Journal 101: 36–49.

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(2010). The Hymenophyllaceae of the Pacific area. 2. Hymenophyllum (excluding subgen. Hymenophyllum). Bulletin of the National Museum of Nature and Science, Series B (Botany), Tokyo 36: 43–59.


(2009). New Records of Polyphlebium borbonicum, an African Filmy Fern, in the New World and Polynesia. American Fern Journal 99: 200–206.

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(2009). Hemi-epiphytism in Vandenboschia collariata (Hymenophyllaceae). Brittonia 61: 392–397.

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